1911年春,国家局势动荡,上海召开武术同盟大会，邀请南北武术宗师霍元甲、黄飞鸿莅沪共商大计，但却落入日本人的陷阱之中，霍元甲离奇死亡，被嫁祸给宝芝林弟子闫应元（郭品超饰）、精武门弟子翟枫（何润东饰），两人被迫双双逃亡。经过曲折离奇的遭遇，两人心性改变，走上了不同的人生道路。
日本人的傀儡柯亦难通过侵吞父亲的资产，一跃而为显赫。种种鬼蜮伎俩不断蚕食上海滩上正义力量，锋芒直指精武门、宝芝林！在种种危难局面下，翟枫发誓重振精武门，却要忍受日本人的种种非人折磨，更要面对全上海的鄙夷、非难和打击。而闫应元为了拯救宝芝林，卧底柯家，眼睁睁看着宝芝林一步一步滑向深渊，心在流血却还要忍受全天下的误会，尤其是爱侣反目，同志成仇，兄弟相残，闫应元一天天迷失在困境之中；英雄出处，百炼成钢，兄弟二人殊途同归，共同搏击黑云压城的上海滩！
山本调整对策，使出更阴毒手段，利用霍耀堂希冀复辟的心理引诱他出面夺取清室珠宝，目的让精武门自相残杀，彻底毁灭翟枫的斗争意志！经过一场大混战，上海正义武林界遭到毁灭性打击。霍耀堂垂死之际终于醒悟，将女儿林思羽(郝蕾饰)托付给翟枫，含笑而去。日本人与上海三大家族陷入更深的旋涡风暴中！
时局混乱不堪，第一次世界大战爆发，远东局势波谲云诡，山本和后来的日本武术第一高手阪西联合布下天罗地网，妄想夺取德国人的战争之王——福克型飞机……日本人苦心设计，企图扑杀一切敌对势力，闫应元等候已久的机会终于出现，……叵测的命运，看不见的明天，他们能逃出重重网罗吗？！
This is a story hinging on an event that has never been seen. Huo Yuan-jia, northern China’s most celebrated kung fu master, meets Huang Fei-hong, the south’s most revered martial artist from the south. Both men are famed for their mastery of the Mi-zong Step and Shadowless Kick respectively. However, the sudden death of Huo electrifies everyone.
Suspicions are rife that several Japanese despised him and hatched a plot to murder him. Not stopping here, budding martial artist Zhai and Yan, students of Huo and Huang, are driven by vengeful spite and indignation. Smart-but-playful Zhai and gullible-but-gifted Yan need to overcome their own weakness in kung fu so that they can exact revenge.
Enveloped within the political turmoil in 1910s, they also stand for the rights of the oppressed within a country long plagued by rapacious foreign powers, paving the way to a heart-wrenching conclusion.
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